GCA_000020025.1 Npun_F1315 definition 3-phosphoshikimate 1-carboxyvinyltransferase GCA_000020025.1 Npun_F3918 definition shikimate/quinate 5-dehydrogenase GCA_000020025.1 Npun_R4233 definition shikimate 5-dehydrogenase GCA_000020025.1 Npun_R6293 definition shikimate kinase