GCA_000018585.1 P9211_00141 definition tRNA-dihydrouridine synthase GCA_000018585.1 P9211_11041 definition 16S rRNA uridine-516 pseudouridylate synthase GCA_000018585.1 P9211_14011 definition Putative tRNA pseudouridine 55 synthase GCA_000018585.1 P9211_16531 definition tRNA pseudouridine synthase A